Exoskeleton
An exoskeleton (from Greek έξω éxō "outer"[1] and σκελετός skeletós "skeleton"[2][3]) is a skeleton that is on the exterior of an animal to both support the body shape and protect the internal organs, in contrast to an internal endoskeleton (e.g. that of a human) which is enclosed underneath other soft tissues. Some large, hard and non-flexible protective exoskeletons are known as shell or armour.
"Robot exoskeleton" redirects here. For the type of machine, see powered exoskeleton.Examples of exoskeletons in animals include the cuticle skeletons shared by arthropods (insects, chelicerates, myriapods and crustaceans) and tardigrades, as well as the skeletal cups formed by hardened secretion of stony corals and the prominent mollusc shell shared by snails, clams, tusk shells, chitons and nautilus. Some vertebrate animals, such as the turtle, have both an endoskeleton and a protective exoskeleton.
Role[edit]
Exoskeletons contain rigid and resistant components that fulfil a set of functional roles in addition to structural support in many animals, including protection, respiration, excretion, sensation, feeding and courtship display, and as an osmotic barrier against desiccation in terrestrial organisms. Exoskeletons have roles in defence from parasites and predators and in providing attachment points for musculature.[4]
Arthropod exoskeletons contain chitin; the addition of calcium carbonate makes them harder and stronger, at the price of increased weight.[5] Ingrowths of the arthropod exoskeleton known as apodemes serve as attachment sites for muscles. These structures are composed of chitin and are approximately six times stronger and twice the stiffness of vertebrate tendons. Similar to tendons, apodemes can stretch to store elastic energy for jumping, notably in locusts.[6] Calcium carbonates constitute the shells of molluscs, brachiopods, and some tube-building polychaete worms. Silica forms the exoskeleton in the microscopic diatoms and radiolaria. One mollusc species, the scaly-foot gastropod, even uses the iron sulfides greigite and pyrite.
Some organisms, such as some foraminifera, agglutinate exoskeletons by sticking grains of sand and shell to their exterior. Contrary to a common misconception, echinoderms do not possess an exoskeleton and their test is always contained within a layer of living tissue.
Exoskeletons have evolved independently many times; 18 lineages evolved calcified exoskeletons alone.[7] Further, other lineages have produced tough outer coatings, such as some mammals, that are analogous to an exoskeleton. This coating is constructed from bone in the armadillo, and hair in the pangolin. The armour of reptiles like turtles and dinosaurs like Ankylosaurs is constructed of bone; crocodiles have bony scutes and horny scales.