
List of longest-living organisms
This is a list of the longest-living biological organisms: the individual(s) (or in some instances, clones) of a species with the longest natural maximum life spans. For a given species, such a designation may include:
For the oldest known life forms, see Earliest known life forms.
The definition of "longest-living" used in this article considers only the observed or estimated length of an individual organism's natural lifespan – that is, the duration of time between its birth or conception, or the earliest emergence of its identity as an individual organism, and its death – and does not consider other conceivable interpretations of "longest-living", such as the length of time between the earliest appearance of a species in the fossil record and the present (the historical "age" of the species as a whole), the time between a species' first speciation and its extinction (the phylogenetic "lifespan" of the species), or the range of possible lifespans of a species' individuals. This list includes long-lived organisms that are currently still alive as well as those that are dead.
Determining the length of an organism's natural lifespan is complicated by many problems of definition and interpretation, as well as by practical difficulties in reliably measuring age, particularly for extremely old organisms and for those that reproduce by asexual cloning. In many cases the ages listed below are estimates based on observed present-day growth rates, which may differ significantly from the growth rates experienced thousands of years ago. Identifying the longest-living organisms also depends on defining what constitutes an "individual" organism, which can be problematic, since many asexual organisms and clonal colonies defy one or both of the traditional colloquial definitions of individuality (having a distinct genotype and having an independent, physically separate body). Additionally, some organisms maintain the capability to reproduce through very long periods of metabolic dormancy, during which they may not be considered "alive" by certain definitions but nonetheless can resume normal metabolism afterward; it is unclear whether the dormant periods should be counted as part of the organism's lifespan.
List of longest-living organisms[edit]
Microorganisms[edit]
Some endoliths have extremely long lives. In August 2013, researchers reported evidence of endoliths in the ocean floor, perhaps millions of years old, with a generation time of 10,000 years.[23] These are slowly metabolizing and not in a dormant state. Some Actinomycetota found in Siberia are estimated to be half a million years old.[24][25][26]